Fossil

Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants.The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the absolute ages of rocks and the fossils they host.There are many examples of paleolithic stone knives in Europe, with fossil echinoderms set precisely at the hand grip, dating back to Homo heidelbergensis and Neanderthals.Pliny also makes one of the earlier known references to toadstones, thought until the 18th century to be a magical cure for poison originating in the heads of toads, but which are fossil teeth from Lepidotes, a Cretaceous ray-finned fish.[21] There is no such direct mythological connection known from prehistoric Africa, but there is considerable evidence of tribes there excavating and moving fossils to ceremonial sites, apparently treating them with some reverence.[22] In Japan, fossil shark teeth were associated with the mythical tengu, thought to be the razor-sharp claws of the creature, documented some time after the 8th century AD.One good example is the famous scholar Huang Tingjian of the Song dynasty during the 11th century, who kept a specific seashell fossil with his own poem engraved on it.[23] In his Dream Pool Essays published in 1088, Song dynasty Chinese scholar-official Shen Kuo hypothesized that marine fossils found in a geological stratum of mountains located hundreds of miles from the Pacific Ocean was evidence that a prehistoric seashore had once existed there and shifted over centuries of time.After observing the existence of seashells in mountains, the ancient Greek philosopher Xenophanes (c. 570 – 478 BC) speculated that the world was once inundated in a great flood that buried living creatures in drying mud.[15] These grew into the shepherd's crowns of English folklore, used for decoration and as good luck charms, placed by the doorway of homes and churches.Near the end of his 1796 paper on living and fossil elephants he said: All of these facts, consistent among themselves, and not opposed by any report, seem to me to prove the existence of a world previous to ours, destroyed by some kind of catastrophe.[40] Early naturalists well understood the similarities and differences of living species leading Linnaeus to develop a hierarchical classification system still in use today.When Darwin wrote On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life, the oldest animal fossils were those from the Cambrian Period, now known to be about 540 million years old.The fossil record is life's evolutionary epic that unfolded over four billion years as environmental conditions and genetic potential interacted in accordance with natural selection."[44] Molecular biologists, using phylogenetics, can compare protein amino acid or nucleotide sequence homology (i.e., similarity) to evaluate taxonomy and evolutionary distances among organisms, with limited statistical confidence.Synchrotron X-ray tomographic analysis of early Cambrian bilaterian embryonic microfossils yielded new insights of metazoan evolution at its earliest stages.Fossils of two enigmatic bilaterians, the worm-like Markuelia and a putative, primitive protostome, Pseudooides, provide a peek at germ layer embryonic development.[jargon] This research is a notable example of how knowledge encoded by the fossil record continues to contribute otherwise unattainable information on the emergence and development of life on Earth.For example, the research suggests Markuelia has closest affinity to priapulid worms, and is adjacent to the evolutionary branching of Priapulida, Nematoda and Arthropoda.[86][87][88][89] In 2014, Mary Schweitzer and her colleagues reported the presence of iron particles (goethite-aFeO(OH)) associated with soft tissues recovered from dinosaur fossils.Microfossils may either be complete (or near-complete) organisms in themselves (such as the marine plankters foraminifera and coccolithophores) or component parts (such as small teeth or spores) of larger animals or plants.Microfossils are of critical importance as a reservoir of paleoclimate information, and are also commonly used by biostratigraphers to assist in the correlation of rock units.Fossil resin (colloquially called amber) is a natural polymer found in many types of strata throughout the world, even the Arctic.[104] The main importance of subfossil vs. fossil remains is that the former contain organic material, which can be used for radiocarbon dating or extraction and sequencing of DNA, protein, or other biomolecules.[12][13]Stromatolites are layered accretionary structures formed in shallow water by the trapping, binding and cementation of sedimentary grains by biofilms of microorganisms, especially cyanobacteria.The most widely supported explanation is that stromatolite builders fell victims to grazing creatures (the Cambrian substrate revolution), implying that sufficiently complex organisms were common over 1 billion years ago.[118] While prokaryotic cyanobacteria themselves reproduce asexually through cell division, they were instrumental in priming the environment for the evolutionary development of more complex eukaryotic organisms.Cyanobacteria (as well as extremophile Gammaproteobacteria) are thought to be largely responsible for increasing the amount of oxygen in the primeval Earth's atmosphere through their continuing photosynthesis.In modern microbial mats, debris from the surrounding habitat can become trapped within the mucus, which can be cemented by the calcium carbonate to grow thin laminations of limestone.[121][122] According to one hypothesis, a Corinthian vase from the 6th century BCE is the oldest artistic record of a vertebrate fossil, perhaps a Miocene giraffe combined with elements from other species.[125] However, a subsequent study using artificial intelligence and expert evaluations reject this idea, because mammals do not have the eye bones shown in the painted monster.
Montage of multiple fossils. Clockwise from top left: Onychocrinus and Palaeosinopa ; bottom row: Gryphaea and Harpactocarcinus
Ceratopsian skulls are common in the Dzungarian Gate mountain pass in Asia, an area once famous for gold mines, as well as its endlessly cold winds. This has been attributed to legends of both gryphons and the land of Hyperborea.
Fossil shells from the cretaceous era sea urchin, Micraster , were used in medieval times as both shepherd's crowns to protect houses, and as painted fairy loaves by bakers to bring luck to their bread-making.
Georges Cuvier's 1812 skeletal reconstruction of Anoplotherium commune based on fossil remains of the extinct artiodactyl from Montmartre in Paris, France
Ichthyosaurus and Plesiosaurus from the 1834 Czech edition of Cuvier 's Discours sur les revolutions de la surface du globe
Phacopid trilobite Eldredgeops rana crassituberculata . The genus is named after Niles Eldredge
Stratigraphy of the Montañita-Olón locality of the Dos Bocas Formation . Stratigraphy is a useful branch when it comes to the understanding of the successive layers of rock and their fossiliferous content, giving insight into the relative age of fossils
Permineralized bryozoan from the Devonian of Wisconsin
The star-shaped holes ( Catellocaula vallata ) in this Upper Ordovician bryozoan represent a soft-bodied organism preserved by bioimmuration in the bryozoan skeleton. [ 92 ]
Examples of index fossils
Microfossils about 1 mm
The wasp Leptofoenus pittfieldae trapped in Dominican amber , from 20 to 16 million years ago. It is known only from this specimen.
Eroded Jurassic plesiosaur vertebral centrum found in the Lower Cretaceous Faringdon Sponge Gravels in Faringdon, England. An example of a remanié fossil.
A subfossil dodo skeleton
Lower Proterozoic stromatolites from Bolivia , South America
An example of a pseudofossil: Manganese dendrites on a limestone bedding plane from Solnhofen , Germany; scale in mm
Dinosaur BonesFossil (disambiguation)Fossilization (linguistics)OnychocrinusPalaeosinopaGryphaeaHarpactocarcinusClassical Latingeological ageshellsexoskeletonsmicrobespetrified woodTiktaalikCanadaPaleontologyevolutionarystratageological timescalerelative agesradiometric datingabsolute agespermineralizationauthigenic mineralizationcarbonizationmicrometredinosaursmineralizedvertebrateschitinouscalcareousinvertebratesanimal trackscoprolitestrace fossilsbiochemicalbiosignaturesTrace fossilMicrofossilFossil preparationIndex fossilList of fossilsList of fossil sitesLagerstätte fossil bedsList of transitional fossilsList of human evolution fossilsBiogeographyExtinction eventGeochronologyGeologic time scaleGeologic recordHistory of lifeOrigin of lifePaleoclimatologyTimeline of evolutionTransitional fossilAvian flightMulticellsFlagellaMammalian auditory ossiclesMosaic evolutionNervous systemsButterfliesCephalopodsCetaceansFishesHumansInsectsMammalsMolluscsPlantsReptilesSea cowsSpidersIntroduction to evolutionCommon descentPhylogenyCladisticsBiological classificationHistory of paleontologyTimeline of paleontologyBiostratigraphyIchnologyInvertebrate paleontologyMicropaleontologyMolecular paleontologyPalaeoxylologyPaleobiologyPaleobotanyPaleoecologyPaleogeneticsPaleolimnologyPaleomycologyPaleoneurobiologyPaleopathologyPaleopedologyPaleotempestologyPaleozoologyPalynologySclerochronologyTaphonomyVertebrate paleontologyevolutionhistory of life on EarthPaleontologistsspeciesDzungarian GatepaleolithicechinodermsHomo heidelbergensisNeanderthalshippopotamuscretaceousMicrasterHerodotosHyperboreagryphonsProtoceratopsAristotlevaporousexhalationsPersianAvicennapetrifyingfluidsAlbert of SaxonynaturalistsPliny the Eldertongue stonesglossopetrahorns of AmmonammonitestoadstonesLepidotesPlains tribesthunderbirdHomo erectusdragonaphrodisiacsHuang TingjianSong dynastyDream Pool Essaysscholar-officialShen Kuogeological stratumPacific Oceanshifted over centuries of timepetrifiedbamboosYan'anShaanxiclimate changeChristendomNoah's Arkancient Greek philosopherXenophanesThe Book of HealingDeinotherium giganteumCyclopesGreek mythologyelephantNorse mythologySuffolkAnoplotheriumartiodactylMontmartreRenaissanceLeonardo da VinciIchthyosaurusPlesiosaurusCuvierNicholas StenoRobert HookemicrographsMicrographiaforamsWilliam Smith (1769–1839)law of superpositionGeorges CuviercatastrophismMary AnningichthyosaurLinnaeusOn the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for LifeCambrianmicrofossilsEdiacara biotaeukaryotesgeologyrelative agestratigraphersabsolute datingpotassium/argonargon/argonuranium seriesradiocarbon datingevolutionary biologyphylogeneticsamino acidnucleotidePhacopidtrilobiteNiles EldredgePhacopsDevonianStephen Jay Gouldpunctuated equilibriumSynchrotrontomographicembryonicmetazoanMarkueliaprotostomegerm layerarthropodSiberiadiageneticPriapulidaNematodaArthropodaRelative datingDos Bocas Formationstratigraphysedimentaryfaultserosionindex fossilscontinentsmutationsmolecular clocksCambrian explosionGhost lineageSignor–Lipps effectsoft-bodied organismsmollusksbrachiopodsarthropodsLagerstätteLagerstättenanoxicpresentMaotianshan ShalesBurgess ShaleHunsrück SlatesJurassicSolnhofen LimestoneCarboniferousMazon CreekaragonitecalciteFort Drum, FloridaBusyconscleractinianIsraelpentameridIndianaBird Spring Formationbryozoangroundwatercell wallplant cellfeathersdiagenesiscalciumphosphatemineralssulfurpyritePrecambrianPleuroceras solareTriarthrus eatoniHunsrück SlateLudlowville Formationsilicificationsilica dehydratequartzchalcedonyElimia teneraostracodsgastropodsDeccan TrapsFloridaQueenslandPlesiosaurSouth Australian MuseumendocastbivalveWaynesville FormationAustriaAviculopecten subcardiformisLogan FormationsideriteMazon Creek fossil bedsCompression fossilscarbonaceous filmcycloneuralianWaukesha BiotalycopodWisconsinMary SchweitzergoethitehaemoglobinchelationcollagensessileoystersubstratesclerobiontsOrdoviciangeologic periodssedimentsearthwormsWilliam BucklandbezoarRusophycusSaskatchewanClimactichnitesmolluskProtists in the fossil record"macro" fossilsforaminiferacoccolithophoressporespaleoclimatebiostratigraphersLeptofoenus pittfieldaeDominican amberpolymerArcticTriassicCenozoicadaptationvertebrateZombie taxonFossil woodbotanical nameAraucarioxylonAraucariaPalmoxylonchinkapinfecal depositssequencing of DNAproteinisotopeweatheringorganismsMesozoicQuaternarychironomidostracodcarapacesdiatomsTheba geminataseashellsHolocenePleistoceneBiosignaturefossil fuelsMolecular fossilszirconsStromatoliteEarliest known life formsstromatolitesBoliviaaccretionarystructuresbiofilmsmicroorganismscyanobacteriaArcheancoloniesProterozoicprimordialeukaryotechlorophytesgreen algaeColleniaCambrian substrate revolutionevolutionary radiationend-Ordovicianend-Permian extinctionsprokaryoticevolutionary developmenteukaryoticextremophileGammaproteobacteriaoxygenatmospherephotosynthesiscarbon dioxidesunlightlimestoneoncolitesfossil recordThrombolitesNamibiaSolnhofenPseudofossilsdendritemoss agatesConcretionsdinosaurbiomineralsextraterrestrial lifeorganic componentsbioticCuriosityOpportunityroversbiosphereautotrophicchemotrophicchemolithoautotrophicfluvio-lacustrine environmentsplainsrivershabitablehabitabilityorganic carbonFossil tradingFossil collectingtraditional medicinetalismansplacebo effectantacidessential mineralsTraditional Chinese medicineRuyang CountyDhankar GompaammoniteEoceneGreen River FormationAsaphus kowalewskiiMegalodonCarcharodontosaurusSahara DesertshrimpPetrified Forest National ParkAraucaria mirabilisPatagoniaArgentinaJurassic PeriodgastropodPlioceneCyprusserpulid wormSilurianOrthocerasechinoidbrachiopodGuadalupianPermianAgatizedHawthorn GroupOligoceneMioceneBaltic SeaGotlandTorotoro National ParkBioerosionCryptosporeEndolithList of fossil parksLiving fossilSchultz's ruleShark toothBibcodeNatureNoffke, NoraAstrobiologyExciteMindspark Interactive NetworkAssociated PressMeteorologyUniversity of Chicago PressPeople's Daily NetCambridge University PressSingapore University PressNational Geographic Societyda Vinci, LeonardoDarwin, CharlesThe Origin of SpeciesFortey, RichardSedimentary GeologyProthero, Donald R.Columbia University PressTalkOrigins ArchiveDictionary.comNature GeoscienceMars Exploration Program Analysis GroupScienceCiteSeerXHellenic Journal of GeosciencesNational GeographicKhan AcademyNatural History Museum, LondonAtlas ObscuraWayback MachineOlivia JudsonThe New York TimesIn Our TimeErnest IngersollEncyclopedia AmericanaNew International EncyclopediaSkeletonizationStages of human developmentCell deathNecrosisAvascular necrosisCoagulative necrosisLiquefactive necrosisGangrenous necrosisCaseous necrosisFat necrosisFibrinoid necrosisTemporal lobe necrosisProgrammed cell deathAnoikisApoptosisAutophagyIntrinsic apoptosisNecroptosisParaptosisParthanatosPhenoptosisPseudoapoptosisPyroptosisAutolysisAutoschizisEscharImmunogenic cell deathIschemic cell deathPyknosisKaryorrhexisKaryolysisMitotic catastropheSuicide geneAbortionAccidental deathAutopsyBrain deathBrainstem deathClinical deathDeath by natural causesDeath rattleDysthanasiaEnd-of-life careEuthanasiaLazarus signLazarus syndromeOrgan donationTerminal illnessUnnatural deathCauses of death by rateNotable deaths by yearExpressions related to deathNatural disastersPeople by cause of deathPremature obituariesPreventable causes of deathUnusual deathsList of ways people dishonor the deadList of ways people honor the deadBirthday effectChild mortalityExcess mortalityGompertz–Makeham law of mortalityInfant mortalityKaroshiMaternal deathMaternal mortality in fictionMemento moriMicromortMortality rateMortality saliencePerinatal mortalityPallor mortisLivor mortisAlgor mortisRigor mortisPutrefactionDecompositionCryopreservationCryonicsNeuropreservationEmbalmingMacerationMummificationPlastinationProsectionTaxidermyDisposalBurialNatural burialSky burialCremationDismembermentExcarnationPromessionResomationBeating heart cadaverBody donationCadaveric spasmCoffin birthDeath erectionDissectionGibbetingPostmortem caloricityPost-mortem intervalCarrionCemeteryConsciousness after deathAfterlifeTukdamCrematoriumDesecration of gravesEternal oblivionExaminationFuneralInternetMidwifeMourningOnline mourningObituaryReincarnationPalingenesisSaṃsāraResurrectionUnderworldGhostsNear-death experienceNear-death studiesNecromancyOut-of-body experienceSéanceAbortion lawAdministrationCapital punishmentCause of deathCivil deathCoronerDeath-qualified juryDeath certificateDeclared death in absentiaDeath rowDying declarationInquestLegal deathMurderNecropoliticsProhibition of dyingRight to dieSuspicious deathTrust lawForensic pathologyFuneral directorMortuary scienceNecrobiologyPost-mortem chemistryPost-mortem photographyBiostratinomyThanatologyApparent deathChinese burial moneyCoins for the deadDark tourismDarwin AwardsDeath and cultureDeath anniversaryDeath anxietyDeath deityPersonification of deathDying-and-rising godPsychopompDeath campDeath driveDeath educationDeath from laughterDeath hoaxDeath knellDeath marchDeath messengerDeath notificationDeath panelDeath poemDeath poseDeath-positive movementDeath squadDeath threatDeath trajectoryDignified deathExtinctionFestival of the DeadFascination with deathHierarchy of deathHomicideImmortalityLast ritesLongevityMartyrMuseum of DeathNecronymNecrophiliaNecrophobiaPhilosophy of deathPredationSacrificeSuicideAssisted suicideThanatosensitivityOutline