Oryzomyini
[2] The genera Holochilus (including Lundomys at the time), Pseudoryzomys, and Zygodontomys were not included at the time because of their tetralophodont molars (lacking complete mesoloph(id)s); instead, Holochilus was considered to be a sigmodont, related to Sigmodon and Reithrodon, and Pseudoryzomys and Zygodontomys were considered to be members of Phyllotini, another large South American tribe.[2] Although lacking mesoloph(id)s, these genera share other characters with oryzomyines, and a series of papers by Robert Voss and coworkers in the early 1990s established their membership in Oryzomyini.Megaoryzomys, an extinct giant rat from the Galápagos Islands, has been allocated to both Oryzomyini and Thomasomyini, but its correct classification currently remains unclear.[17] In the 2000s, Marcelo Weksler published several studies in which he used evidence from IRBP, a nuclear gene, and morphology to assess the relationships among the members of Oryzomyini.[36] Oryzomyines range from New Jersey in the north, where the Marsh Rice Rat (Oryzomys palustris) is found, to Tierra del Fuego in the south, where Oligoryzomys magellanicus occurs.Extinct species are known from Jamaica (Oryzomys antillarum), the Galápagos Islands (Nesoryzomys and Aegialomys galapagoensis), Fernando de Noronha (Noronhomys), and the Lesser Antilles north to Anguilla (Megalomys, Oligoryzomys victus, and several unidentified genera).